The Prime Minister has finished and he can take it from me that he’s finished.
Mr Bercow - a Tory MP before being elected Speaker in 2009 - stood up and gestured for the Prime Minster to sit down as he was answering a question in the Commons.
Mr Cameron could be heard to say: "I haven’t finished!"
After an avalanche of jeering from other MPs, Mr Bercow told Mr Cameron: "The Prime Minister has finished and he can take it from me that he’s finished."
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In a dig at Mr Bercow - believed to be 5ft 6in tall - Mr Cameron said that health minister Simon Burns's driver had accidentally hit the Speaker's car while reversing.
According to Mr Cameron's joke, the Speaker told Mr Burns he wasn't "happy", to which Mr Burns replied: "So which one are you?" - a reference to Snow White and Seven Dwarfs.
In February, Mr Bercow wrote to all three party leaders asking them for ideas how to tone down the noise during Prime Minister's Questions.
At the time, Tory MP Alec Shelbrooke put the blame on the disorder at PMQs on Mr Bercow, accusing him of being "biased".
He said it was his fault that he was unable to control behaviour at PMQs, unlike previous speaker Baroness Betty Boothroyd.
The Elmet and Rothwell MP wrote on Twitter: "Bercow needs to look in the mirror. Betty never had the need to resort to whining. His biased approach is why he's lost control of PMQs."
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